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One might also consider, based on some passages in Morgoth's Ring, the role of corrupted Elvish fear- whether those of Avari who had already turned to the Darkness before death, or those souls which refused the summons to Mandos. Tolkien treats these as malign "spirits" haunting the shadows, with whom the living should never have ado (those who do so are "the necromancers, and of the train of Sauron.") Could not some of these 'powered' the werewolves?
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The entire plot of The Lord of the Rings could be said to turn on what Sauron didn’t know, and when he didn’t know it.
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