The Two Towers are Orthanc and Minas Morgul. Not totally sure why they were specifically picked out....Orthanc is obvious but either Morgul, Cirith Ungol or Barad-Dur could have been the other one. For proof look at the cover of the 1997 paperback which like the other 2 has original Tolkien drawings on the cover.
In more detail for those who don't have it the cover shows Minas Morgul on the left and and Orthanc on the right. Below Minas Morgul there is a Crescent Moon hinting at former name of Minas Ithil..below Orthanc is the white hand of Saruman. Between them lies Mt Doom, the one ring and a winged Nazgul.
PC.
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Ugluk u bagronk sha pushdug Saruman-glob bubhosh skai'...
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