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When I read the books I get the strong impression that it is Orthanc and Barad-Dûr. A large part of The Two Towers focuses mainly on Orthanc. When Orthanc is defeated their attention is turned to the bigger threat, Barad-Dûr.
There is a comparison going on between Saruman and Sauron. Saruman is trying to immitate Barad-Dûr with Orthanc, but he can not compare. Though Saruman was the main focus of most of The Two Towers and there is a definate feeling of victory when he is defeated, Tolkien soon reminds us that this was just the first round. There is still a greater enemy (second tower) to think of. Thus The Two Towers not Orthanc or Saruman's Tower.
Anyway that was my interpretation.
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If you would convince a man that he does wrong, do right. Men will believe what they see.~Henry David Thoreau
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